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BEIRUT — Syrian government forces backed by heavy Russian airstrikes have seized three neighborhoods inside Palmyra, a town with famed Roman-era ruins that fell to the Islamic State group last May, state media reported Saturday.

Russian jets carried out 40 air sorties near Palmyra in the past day, hitting 158 targets and killing over 100 militants, Russia’s defense ministry said.

Syrian troops and allied militiamen have taken up positions in the three neighborhoods that are part of the modern town, according to the Syrian Observatory for Human Rights, a Britain-based monitoring group.

Palmyra, affectionately known as the “bride of the desert,” used to attract tens of thousands of tourists every year. The Islamic State drove out government forces in a matter of days and later demolished some of the best-known monuments in the UNESCO world heritage site. The extremists believe ancient ruins promote idolatry. The militants also demolished the town’s infamous Tadmur prison, where thousands of Syrian government opponents were reported to have been tortured.

Retaking the town would be a major victory for President Bashar Assad’s government, which has made steady gains in recent months against the Islamic State and other insurgents. Syria’s government has been assisted in large part by Russia’s air campaign. Earlier this month, Russian President Vladimir Putin ordered the surprise pullout of some Russian warplanes from Syria, but said that strikes against the Islamic State group and the al-Qaeda-linked Nusra Front will continue. Those groups have been excluded from a Russian- and U.S.-brokered cease-fire that began on Feb. 27 and has largely held.

The battle for Palmyra, now entering its fourth week according to the Observatory, has not been easy. Government forces lost at least 18 soldiers on Friday alone, including a major general, the Observatory and Islamic State-affiliated media sites reported. The Observatory’s figures indicate at least 56 soldiers have died in fighting this week.

Footage broadcast on Lebanese stations aligned with the Syrian government showed smoke rising over Palmyra’s skyline, as tanks and helicopters fired at positions inside the town. The Islamic State began evacuating civilians this week to other parts of its territories in Syria. No civilians remain in the town, a Palmyra resident who left last week told The Associated Press.

The fate of the archaeological site was not immediately clear. Activists citing sources among advancing government forces said the two sides were fighting over the area, while the Observatory said government forces had retaken the site. Syrian state media made no mention of the area.

The head of Syria’s antiquities and museums authority described the condition of the remaining monuments — including the grand amphitheater, the market place, public baths and a long colonnade — as “excellent,” with only minor damage, based on footage captured by a drone and broadcast on Russian television Friday.

Maamoun Abdulkarim, head of the government’s antiquities and museums department, said civil servants are prepared to assess the monuments and plan for their restoration “as soon as we are given the green light from the Syrian army.”

“We will present our studies to UNESCO so that Palmyra can once again be a world heritage site,” said Abdulkarim. “Palmyra is not just for Syrians, it is for the world.”

If Syrian government forces retake Palmyra, they will be positioned to advance on the two largest Syrian cities held by the Islamic State, Deir Ezzor and Raqqa. The Islamic State group is on the back foot in Iraq and Syria, where forces on the ground have been backed by US-led airstrikes against the extremists. The U.S.-led international coalition estimates that the group has lost 40 percent of the territory it once held in Iraq and around 20 percent of its territory in Syria.

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